Born at Fairfield Road, Bristol, on 1 October 1865, the eldest child of David Rayner, wholesale textile merchant, and his wife, Edith Fryer. He was educated at Observatory House School, Bristol, under Dr Cooper, and at the Bristol Medical School. After working in the physiological laboratories, he d

Born at Honingham, Norfolk in 1869 he was educated at Bracondale School, Norwich and King's College Hospital, which he entered with an exhibition, and won prizes and scholarships each year from 1887 to 1890. He qualified with the Conjoint Diploma and the London MB in 1892, took the BS in 1894, and g