Born 14 December 1878, the second son and third child of Robert Harry Hughes, MB, who was in general practice at Plymouth, and his wife Laetitia Cavanagh Jervis. His elder brother Lawson Jervis Hughes, after training at Guy's Hospital, was killed in the South African war, when he was serving as a mo

Born 10 March 1881 at Hinckley, Leicestershire, the second child and only son of Edward Simpson, a schoolmaster, and Charlotte McCann, his wife. He was educated at Mill Hill School and St John's College, Cambridge, where he was a scholar, and took first-class honours in the Natural Sciences Tripos,